Carolinea 77

148 Carolinea 77 (2019) to each bothridium. Two further very large paired sigillary fields occur in the central region of the prodorsum. Distinct centroprodorsal groove ab- sent. Dorsosejugal furrow deep and fairly wide. Notogaster (Figs 1, 3) Notogaster shield-like, covers most of hystero- soma. Observed from a viewpoint perpendicular to the circumgastric scissure, it is slightly oval, about 1,16 times as long as wide. Notogastral length 402 (370-431) µm, width 347 (315-368)µm. In lateral perspective notogaster hemispherical, robust and fairly flat, maximum height about 156 µm, corresponding to 0,4 times the notogastral length. Dorsal notogastral margin in lateral view almost evenly rounded, only anteriorly slightly flattened. Area porosae between setae h1 ab- sent. Spinae adnatae absent. Eleven pairs of notogastral setae present. Setae of c-, l-, and h- series slender, smooth, curved in lateral aspect, approximately 55-68 µm long; their alveoli arranged along two symmetrical lon- gitudinal rows which are arched, in dorsal view. In well cleared notogasters a thin, brown line is visible paraxial to the insertion points of setae c2 to h2. Anterior to seta c2 the lines become in- distinct. Posteriorly the lines are delimited by the oval sigillary field between setae h1 and ps1. Us- ing only a light microscope and without employ- ing histological methods, it was not possible to conclusively identify the exact nature of the linear structure and the region immediately medial to it. One hypothesis is that the line represents a shal- low longitudinal groove, which forms the border of a lightly sclerotized strip or very low ridge on which the insertion points of setae c2 to h1 are located, with all of the alveoli being positioned paraxial to the groove. Notogastral setae darkly pigmented except for the short proximal base part which is lighter in colour and also thinner than the adjoining more distal portion. Mutual distance between setae c1: 25 µm, between setae lm: 174 µm. Distance be- tween setal pair c2 about 4 times that of c1. A diminutive lightly coloured spot present postero- mediad the insertion point of each seta la and lp. In dorsal perspective setae c1 directed towards the anterior, the remainder, c2-h3, radially direct- ed. Pseudanal setae ps1, ps2, ps3 marginally positioned on posterior ventrolateral notogastral border, thin, smooth, finely attenuate and flagel- late; ps1 very long (80-110 µm). Sigillae along lateral and posterolateral margins of the notogaster form two long bands, ending an- teriorad just posterior to insertion points of setae c2. Scattered but more or less regularly arranged sigillae also anteriorly in area between insertions of setae c1 and c2 and similarly posteriorly in the sector demarcated by insertions of setal pairs h1 and ps1. Lyrifissures ia, im, ip, ih and ips present, with latter two located on ventral border. Opening of latero-opisthonotal gland located midlaterally on notogastral margin between lyrifissure im and ip. Distance ia - im > im - ip. Epimeral region (Figs 2, 3) The epimeral region consists of the contiguous epimeres I-IV, sclerotized coxal fields, each of which is associated with a pair of legs. Tubercle Sa of parastigmatic enantiophyses long, cerati- form, laterally directed and with a sharply poin- ted apex. Sp much shorter, subtriangular. Length of Sa 2,5-3x the length of Sp when observed in ventral perspective. Discidium elongate, conical to subtriangular, acute. On opposite sides of the ventrosejugal groove lie conspicuous, large tu- bercles Va and Vp of ventrosejugal enantiophy- sis, their tips rounded to subtriangular. Tubercle E2p of propodoventral enantiophysis slightly less well defined. E2a very indistinct, represented by a slightly thickened ridge. Formula for epimeral setation: 3-1-3-4. Epimeral setae 1a, 2a short (24-30 µm), the remainder measuring 38-57 µm in length. Setae 1c, 3b, 3c, 4d smooth; all other epimeral setae with double rows of minute rela- tively widely spaced barbs. Seta 3b associated with tubercle Vp. Epimeres I and II have distinct areas of muscle scars, whereas the sigillary fields of epimeres III and IV are joined together into one large area. Significant alveolar or verru- cose sculpturing absent. Anogenital region (Figs 2, 3) Adanal setae ad1, ad2, ad3 smooth, flagellate, only slightly longer (48-63 µm) than the epimeral setae. Their insertion points form a straight line when seen in ventral perspective with seta ad3 being significantly laterally situated. Ratios of the distances of the insertion points of setae ad1, ad2, ad3 to the border of the anal plate are about 1: 2: 4.5 respectively. Genital and anal plates lar- ge and positioned close together. Anal opening distinctly longer and slightly wider than genital opening. Paired genital valves each with a row of six barbed slender setae, g1-g4 inserted close together, g5, g6 more distantly situated. Seta g1 slightly longer than the others. Posterior medial region of genital valves with sigillae. Paired anal