Dr. Julien Kimmig

Leitung Referat Paläontologie und Evolutionsforschung; Kurator

Telefon: +49 721 175-2117



  • Leiter Referat Paläontologie und Evolutionsforschung
  • Kurator: Betreuung der Sammlungen des Referats, Sichtung und Annahme von Sammlungen, Bewertung und Digitalisierung von Sammlungsobjekten, Leihverkehr, Konzeption von Ausstellungen, Durchführung geologisch-paläontologischer Grabungen, etc.
  • Öffentlichkeitsarbeit: Wissenschafts-Twitter, Interviews für Zeitungen und im TV, Beratung Wissenschaftsdokumentationen und Buchprojekte, etc.
  • ...



  • Dozent für Fundamentals of Human Energy Use (EARTH 10) an The Pennsylvania State University
  • Gutachter für nationale und internationale Forschungsartikel und Forschungsanträge
  • Mitglied der Paleontological Society Employment of Paleontologists Ad hoc committee
  •  ...



  • seit 2022: Leitung des Referats Paläontologie und Evolutionsforschung am SMNK

  • seit 2021 Adjunct Assistant Teaching Professor an The Pennsylvania State University

  • 2020 bis 2022 Assistant Research Professor an The Pennsylvania State University

  • 2019 bis 2022 Research Affiliate an der University of Kansas

  • 2016 bis 2019 Collections Manager an der University of Kansas

  • 2014 bis 2016 Postdoctoral Researcher an der University of Saskatchewan

  • 2011 bis 2014 Teaching Assistant an der University of Saskatchewan

  • 2011 bis 2014 2014 Ph.D. Geologie University of Saskatchewan

  • 2009 bis 2011 Adjunct Researcher am Natural History Museum London

  • 2008 bis 2009 2009 M.Sc. Biologie Imperial College London & Natural Historyy Museum London

  • 2007 bis 2008 Museum and Research Assistant am Geowissenschaftliches Museum Göttingen

  • 2005 bis 2008 2008 B.Sc. Geowissenschaften Georg-August Universität Göttingen



ORCID: https://orcid.org/0000-0001-8032-4272

Google Scholar: https://scholar.google.com/citations?user=5fDD9bEAAAAJ&hl=en&oi=ao


Peer reviewed

(*Beaufsichtigter Bachelor Student; **Beaufsichtigter Master Student, #Korrespondierender Autor)


#Kimmig, Julien, Stephen Pates, Rhiannon J. LaVine, L. J. Krumenacker, Anna F. Whitaker, Luke C. Strotz, Paul G. Jamison, Val G. Gunther, Glade Gunther, Matt Witte, Allison C. Daley, Bruce S. Lieberman (2023) New soft-bodied panarthropods from diverse Spence Shale (Cambrian; Miaolingian; Wuliuan) depositional environments. Journal of Paleontology, DOI: 10.1017/jpa.2023.24


#Kimmig, Julien, Rhiannon J. LaVine, James D. Schiffbauer, Sven O. Egenhoff, Kevin L. Shelton & Wade W. Leibach (2023) Annelids from the Cambrian (Wuliuan Stage, Miaolingian) Spence Shale Lagerstätte of northern Utah, USA. Historical Biology, DOI: 10.1080/08912963.2023.2196685


#Bicknell, Russell D.C., Patrick M. Smith & Julien Kimmig (2023) Novel coprolitic records from the Silurian (Přídolí) Wallace Shale of New South Wales. Alcheringa: An Australasian Journal of Palaeontology 47: 24–30. DOI: 10.1080/03115518.2023.2185679


**,#Whitaker, Anna F., James D. Schiffbauer, Derek E.G. Briggs, *Wade W. Leibach & Julien Kimmig (2022) Preservation of soft-bodied fossils and the occurrence of phosphate-associated rare earth elements in the Cambrian (Wuliuan) Spence Shale Lagerstätte. Palaeogeography, Palaeoclimatology, Palaeoecology 592: 110909. DOI: 10.1016/j.palaeo.2022.110909


#Bicknell, Russell D. C., Julien Kimmig, Graham E. Budd, David A. Legg, Kenneth S. Bader, Carolin Haug, Dorkas Kaiser, Lukáš Laibl, Jessica N. Tashman & Nicolás E. Campione (2022) Habitat and developmental constraints drove 330 million years of horseshoe crab evolution. Biological Journal of the Linnean Society 136: 155–172. DOI: 10.1093/biolinnean/blab173


#Kimmig, Julien & Brian R. Pratt (2022) Evidence for microbially mediated silver enrichment in a middle Cambrian Burgess Shale-type deposit, Mackenzie Mountains, northwestern Canada. Canadian Journal of Earth Sciences 59: 123–133. DOI: 10.1139/cjes-2021-0035

Article featured in press release.


*,#Leibach, Wade. W, Rudy Lerosey-Aubril, **Anna F. Whitaker, James D. Schiffbauer & Julien Kimmig (2021) First palaeoscolecid from the Cambrian (Miaolingian, Drumian) Marjum Formation of western Utah. Acta Palaeontologica Polonica 66: 663–678. DOI: 10.4202/app.00875.2021

Article featured in press release.


#Yang, Xianfeng, #Julien Kimmig, Dayou Zhai, Yu Liu, Sara R. Kimmig & Shanchi Peng (2021) A juvenile-rich palaeocommunity of the lower Cambrian Chengjiang biota: evidence for palaeo-boom or palaeo-bust? Nature Ecology & Evolution 5: 1082–1088. DOI: 10.1038/s41559-021-01490-4

Cover article.

Article featured in press release.


#Farrell, Úna C., and 103 authors including Julien Kimmig (2021) The Sedimentary Geochemistry and Paleoenvironments Project. Geobiology 19: 545–556. DOI: 10.1111/gbi.12462


#Kimmig, Julien & Paul A. Selden (2021) A new shell-bearing organism from the Cambrian Spence Shale of Utah. Palaeoworld 30: 220–228. DOI: 10.1016/j.palwor.2020.05.003

Article featured in press release.


*,#Leibach, Wade. W, *Nick Rose, Kenneth Bader, Laura J. Mohr, *Kristopher Super & Julien Kimmig (2021) Horseshoe crab trace fossils and associated ichnofauna of the Pony Creek Shale Lagerstätte, Upper Pennsylvanian, Kansas, USA. Ichnos 28: 34–45. DOI: 10.1080/10420940.2020.1811268


#Bicknell, Russell D. C., Patrick Smith, Natalie Schroeder & Julien Kimmig (2020) Reconsidering the ‘phyllocarid’ from the Wade Creek Formation. Alcheringa 44: 481–483. DOI: 10.1080/03115518.2020.1820575


**,#Whitaker, Anna F. & Julien Kimmig (2020) Anthropologically introduced biases in natural history collections, with a case study on the Spence Shale Lagerstätte of Utah. Palaeontologia Electronica 23: a58. DOI: 10.26879/1106


#Lerosey-Aubril, Rudy, Julien Kimmig, Stephen Pates, Jacob Skabelund, Andries Weug & Javier Ortega-Hernández (2020) New exceptionally-preserved panarthropods from the Drumian Wheeler Konservat-Lagerstätte of the House Range of Utah. Papers in Palaeontology 6: 501–531. DOI: 10.1002/spp2.1307


#Yang, Xianfeng, #Julien Kimmig, Bruce S. Lieberman & Shanchi Peng (2020) A new species of the deuterostome Herpetogaster from the early Cambrian Chengjiang biota of South China. The Science of Nature 107: 37. DOI: 10.1007/s00114-020-01695-w


#Kimmig, Julien, *Wade W. Leibach & Bruce S. Lieberman (2020) First occurrence of the problematic vetulicolian Skeemella clavula in the Cambrian Marjum Formation of Utah, USA. Carnets de Géologie 20: 215–221. DOI: 10.4267/2042/70836


**,#Whitaker, Anna, Paul Jamison, James D. Schiffbauer & Julien Kimmig (2020) Re-description of the Spence Shale palaeoscolecids in the light of new morphological features with comments on palaeoscolecid taxonomy and taphonomy. PalZ 94: 661–674­. DOI: 10.1007/s12542-020-00516-9

Article featured in press release.


#Kimmig, Julien, Helena Couto, *Wade W. Leibach & Bruce S. Lieberman (2019) Soft-bodied fossils from the upper Valongo Formation (Middle Ordovician: Dapingian-Darriwilian) of northern Portugal. The Science of Nature 106: 27. DOI: 10.1007/s00114-019-1623-z

Article featured in press release.


#Kimmig, Julien, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff & Bruce S. Lieberman (2019) The Spence Shale Lagerstätte: an Important Window into Cambrian Biodiversity. Journal of the Geological Society of London 176: 609–619. DOI: 10.1144/jgs2018-195

Invited submission.


#Pratt, Brian R. & Julien Kimmig (2019) Extensive burrowing in a middle Cambrian Burgess Shale-type fossil Lagerstätte in northwestern Canada. Geology 47: 231–234. DOI: 10.1130/G45551.1

Article featured in press release.


#Kimmig, Julien, Ronald C. Meyer & Bruce S. Lieberman (2019) Herpetogaster from the early Cambrian of Nevada (Series 2, Stage 4) and its implications for the evolution of deuterostomes. Geological Magazine 156: 172–178. DOI: 10.1017/S0016756818000389


#Seltmann, Katja, and 23 authors including Julien Kimmig (2018) Georeferencing for Research Use (GRU): An integrated geospatial training paradigm for biocollections researchers and data providers. Research Ideas and Outcomes 4: e32449. DOI: 10.3897/rio.4.e32449


#Strotz, Luke C., Erin E. Saupe, Julien Kimmig & Bruce S. Lieberman (2018) Metabolic rates, climate and macroevolution: A case study using Neogene molluscs. Proceedings of the Royal Society B: Biological Sciences 285: 20181292. DOI: 10.1098/rspb.2018.1292

Article featured in news.


#Lieberman, Bruce S. & Julien Kimmig (2018) Museums, Paleontology, and a Biodiversity Science Based Approach. In G. D. Rosenberg, & R. M. Clary (Eds.), Museums at the Forefront of the History and Philosophy of Geology: History Made, History in the Making. Geological Society of America Special Paper 535: 335–348. DOI: 10.1130/20182535(22)


#Strotz, Luke C., Marianna Simões, Matthew Girard, Laura Breitkreuz, Julien Kimmig & Bruce S. Lieberman (2018) Getting somewhere with the Red Queen: Chasing a biologically modern definition of the hypothesis. Biology Letters 14: 20170734. DOI: 10.1098/rslb.2017.0734


#Kimmig, Julien & Brian R. Pratt (2018) Coprolites in the Ravens Throat River Lagerstätte of northwestern Canada: implications for the middle Cambrian food web. Palaios 33: 125–140. DOI: 10.2110/palo.2017.038

Cover article.

Article featured in press release.


#Kimmig, Julien, Luke C. Strotz & Bruce S. Lieberman (2017) The stalked filter feeder Siphusauctum lloydguntheri n. sp. from the middle Cambrian (Series 3: Stage 5) Spence Shale of Utah: Its biological affinities and taphonomy. Journal of Paleontology 91: 902–910. DOI: 10.1017/jpa.2017.57

Article featured in press release.


#Kimmig, Julien & Luke C. Strotz (2017) Coprolites in middle Cambrian (Series 2-3) Burgess Shale-type deposits of Nevada and Utah and their ecological implications. Bulletin of Geosciences 92: 297–309. DOI: 10.3140/bull.geosci.1667


#Lieberman, Bruce S., Richard Kurkewicz, Heather Shinogle, Julien Kimmig & Breandán A. MacGabhann (2017) Disc-shaped fossils resembling porpitids or eldonids from the early Cambrian (Series 2: Stage 4) of western U.S.A. PeerJ 5: e3312. DOI: 10.7717/peerj.3312


#Kimmig, Julien & Brian R. Pratt (2016) Taphonomy of the middle Cambrian (Series 3, Drumian) Ravens Throat River Lagerstätte, Mackenzie Mountains, Northwest Territories, Canada. Lethaia 49: 150–169. DOI: 10.1111/let.12135


#Kimmig, Julien & Brian R. Pratt (2015) Soft-bodied biota from the middle Cambrian (Drumian) Rockslide Formation, Mackenzie Mountains, northwestern Canada. Journal of Paleontology 89: 51–71. DOI: 10.1017/jpa.2014.5


#Kimmig, Julien (2013) Possible secondarily terrestrial lifestyle in the European phytosaur Nicrosaurus kapffi (Late Triassic, Norian) – A preliminary study. In: Triassic Geology and Paleontology: Tanner et al. (Eds.). New Mexico Museum of Natural History and Science, Bulletin 61: 306–312. Link


#Kimmig, Julien & Justin A. Spielmann (2011) Biological factors influencing phytosaur (Archosauria: Phytosauridae) taxonomy: A Prospectus. In: Fossil Record 3: Sullivan et al. (Eds.). New Mexico Museum of Natural History and Science, Bulletin 53: 289–294. Link


#Kimmig, Julien & Gernot Arp (2010) Phytosaur remains from the Norian Arnstadt Formation (Leine Valley, Germany), with reference to European phytosaur habitats. Palaeodiversity 3: 215–224. Link





#Kimmig, Julien (2021) Burgess Shale fauna. The Encyclopedia of Geology, 2nd Edition. Scott Elias & David Alderton (eds.). p. 576–582. DOI: 10.1016/B978-0-12-409548-9.12019-6

Invited submission.


Nicht peer reviewed


Kimmig, Julien (2022) Book Review: Plotnick, R. 2022. Explorers of Deep Time: Paleontologists and the History of Life. The Quarterly Review of Biology 97 (4): 310.


Kimmig, Julien (2017) Book Review: Suttner, T.J. et al. 2016. Planet Earth—In Deep Time. Palaeozoic Series: Devonian and Carboniferous. The Quarterly Review of Biology 92 (3): 306–307.


Kimmig, Julien (2016) Book Review: Gass, K. 2015. Solving the Mystery of the First Animals on Land: The Fossils of Blackberry Hill. Fossil News: Fall 2016: 51.


Kimmig, Julien (2016) Book Review: Penney, D. and J.E. Jepson. 2014. Fossil Insects: An introduction to palaeoentomology. Priscum: Newsletter of the Paleontological Society: Winter 2016: 42–43.


Eingeladene Vorträge


Kimmig, Julien (2023) Das Kambrium – Ursprung des modernen Lebens. Colloquium, Naturwissenschaftlicher Verein Karlsruhe (Germany), April 2023. November 11th, 2023. Seminar talk.


Kimmig, Julien (2022) The middle Cambrian Spence Shale (Miaolingian: Wuliuan) Lagerstätte: Improving our understanding of a key Cambrian ecosystem. The Pennsylvania State University, March 21st, 2022. Seminar talk.


Kimmig, Julien (2021) Understanding Cambrian diversity through time and space. Friedrich-Alexander-Universität Erlangen-Nürnberg, November 22nd, 2021. Seminar talk.


Kimmig, Julien (2019) Cambrian soft-bodied biotas: Understanding Cambrian ecosystems. Gonzaga University, October 18th, 2019. Seminar talk.


Kimmig, Julien, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff & Bruce S. Lieberman (2019) The middle Cambrian Spence Shale (Miaolingian: Wuliuan) Lagerstätte: Improving our understanding of a key Cambrian ecosystem. North American Paleontological Convention, Riverside, June 23rd, 2019. (KEYNOTE)


Kimmig, Julien (2018) The Cambrian: More than just trilobites. The 57th Annual Kansas City Gem and Mineral Show, March 11th, 2018.

Kimmig, Julien (2018) The Cambrian: More than just trilobites. Science on Tap, January 24th, 2018.

Kimmig, Julien (2017) Cambrian Soft-bodied biotas. Kansas and Missouri Paleontological Society, May 28th, 2017.


Kimmig, Julien (2016) Burgess Shale-type deposits: A window into Cambrian paleoecology. Arkansas Tech University, February 9th, 2016. Seminar talk.

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