Dr. Julien Kimmig

Leitung Referat Paläontologie und Evolutionsforschung; Kurator

Telefon: +49 721 175-2117



  • Leiter Referat Paläontologie und Evolutionsforschung
  • Kurator: Betreuung der Sammlungen des Referats, Sichtung und Annahme von Sammlungen, Bewertung und Digitalisierung von Sammlungsobjekten, Leihverkehr, Konzeption von Ausstellungen, Durchführung geologisch-paläontologischer Grabungen, etc.
  • Öffentlichkeitsarbeit: Wissenschafts-Twitter, Interviews für Zeitungen und im TV, Beratung Wissenschaftsdokumentationen und Buchprojekte, etc.
  • ...



  • Dozent für Fundamentals of Human Energy Use (EARTH 10) an The Pennsylvania State University
  • Gutachter für nationale und internationale Forschungsartikel und Forschungsanträge
  • Mitglied der Paleontological Society Employment of Paleontologists Ad hoc committee
  •  ...



  • seit 2022: Leitung des Referats Paläontologie und Evolutionsforschung am SMNK

  • seit 2021 Adjunct Assistant Teaching Professor an The Pennsylvania State University

  • 2020 bis 2022 Assistant Research Professor an The Pennsylvania State University

  • 2019 bis 2022 Research Affiliate an der University of Kansas

  • 2016 bis 2019 Collections Manager an der University of Kansas

  • 2014 bis 2016 Postdoctoral Researcher an der University of Saskatchewan

  • 2011 bis 2014 Teaching Assistant an der University of Saskatchewan

  • 2011 bis 2014 2014 Ph.D. Geologie University of Saskatchewan

  • 2009 bis 2011 Adjunct Researcher am Natural History Museum London

  • 2008 bis 2009 2009 M.Sc. Biologie Imperial College London & Natural Historyy Museum London

  • 2007 bis 2008 Museum and Research Assistant am Geowissenschaftliches Museum Göttingen

  • 2005 bis 2008 2008 B.Sc. Geowissenschaften Georg-August Universität Göttingen



ORCID: https://orcid.org/0000-0001-8032-4272

Google Scholar: https://scholar.google.com/citations?user=5fDD9bEAAAAJ&hl=en&oi=ao


Peer reviewed

(*Beaufsichtigter Bachelor Student; **Beaufsichtigter Master Student, #Korrespondierender Autor)


#Kimmig, Julien & #James D. Schiffbauer (2024) A modern definition of Fossil-Lagerstätten. Trends in Ecology and Evolution 39: . DOI: 10.1016/j.tree.2024.04.004


#Stockey, Richard G., and 55 authors including Julien Kimmig (accepted) Multiple sustained increases in atmospheric oxygen and marine productivity through the Neoproterozoic and Paleozoic Eras. Nature Geoscience 17: . DOI:


#Yang, Xianfeng, #Julien Kimmig, Christopher B. Cameron, Karma Nanglu, Sara R. Kimmig, Danielle de Carle, Caixia Zhang, Mengxiao Yu & Shanchi Peng (2024) An early Cambrian pelago-benthic acorn worm and the origin of the hemichordate larva. Palaeontologia Electronica 27: a17. DOI: 10.26879/1356


#Bicknell, Russell D. C., Julien Kimmig, Patrick M. Smith & Torsten Scherer (2024) An enigmatic euchelicerate from the Mississippian (Serpukhovian) and insights into invertebrate preservation in the Bear Gulch Limestone, Montana, USA. American Museum Novitates 4008: 1–14. DOI: 10.1206/4008.1


#Bicknell, Russell D. C., Jana Bruthansová & Julien Kimmig (2024) Shelly coprolites record durophagous predation in the Late Ordovician Bohdalec Formation (Katian, Prague Basin, Czech Republic). Geobios 82: 1–11. DOI: 10.1016/j.geobios.2023.09.001


#Kimmig, Julien, Rhiannon J. LaVine, James D. Schiffbauer, Sven O. Egenhoff, Kevin L. Shelton & Wade W. Leibach (2024) Annelids from the Cambrian (Wuliuan Stage, Miaolingian) Spence Shale Lagerstätte of northern Utah, USA. Historical Biology 36: 934–943. DOI: 10.1080/08912963.2023.2196685


#Yang, Xianfeng, #Julien Kimmig, James D. Schiffbauer & Shanchi Peng (2023) Herpetogaster collinsi from the Cambrian of China elucidates the dispersal and palaeogeographic distribution of early deuterostomes and the origin of the ambulacrarian larva. PeerJ 11: e16385. DOI: 10.7717/peerj.16385


#Bicknell, Russell D.C. & Julien Kimmig (2023) Clustered and injured Pseudogygites latimarginatus from the late Ordovician Lindsay Formation, Canada. Neues Jahrbuch für Geologie und Paläontologie 309: 199–208. DOI: 10.1127/njgpa/2023/1159


#Kimmig, Julien, Stephen Pates, Rhiannon J. LaVine, L. J. Krumenacker, Anna F. Whitaker, Luke C. Strotz, Paul G. Jamison, Val G. Gunther, Glade Gunther, Matt Witte, Allison C. Daley & Bruce S. Lieberman (2023) New soft-bodied panarthropods from diverse Spence Shale (Cambrian; Miaolingian; Wuliuan) depositional environments. Journal of Paleontology 97: 1025–1048. DOI: 10.1017/jpa.2023.24


#Bicknell, Russell D.C., Patrick M. Smith & Julien Kimmig (2023) Novel coprolitic records from the Silurian (Přídolí) Wallace Shale of New South Wales. Alcheringa: An Australasian Journal of Palaeontology 47: 24–30. DOI: 10.1080/03115518.2023.2185679


**,#Whitaker, Anna F., James D. Schiffbauer, Derek E.G. Briggs, *Wade W. Leibach & Julien Kimmig (2022) Preservation of soft-bodied fossils and the occurrence of phosphate-associated rare earth elements in the Cambrian (Wuliuan) Spence Shale Lagerstätte. Palaeogeography, Palaeoclimatology, Palaeoecology 592: 110909. DOI: 10.1016/j.palaeo.2022.110909


#Bicknell, Russell D. C., Julien Kimmig, Graham E. Budd, David A. Legg, Kenneth S. Bader, Carolin Haug, Dorkas Kaiser, Lukáš Laibl, Jessica N. Tashman & Nicolás E. Campione (2022) Habitat and developmental constraints drove 330 million years of horseshoe crab evolution. Biological Journal of the Linnean Society 136: 155–172. DOI: 10.1093/biolinnean/blab173


#Kimmig, Julien & Brian R. Pratt (2022) Evidence for microbially mediated silver enrichment in a middle Cambrian Burgess Shale-type deposit, Mackenzie Mountains, northwestern Canada. Canadian Journal of Earth Sciences 59: 123–133. DOI: 10.1139/cjes-2021-0035

Article featured in press release.


*,#Leibach, Wade. W, Rudy Lerosey-Aubril, **Anna F. Whitaker, James D. Schiffbauer & Julien Kimmig (2021) First palaeoscolecid from the Cambrian (Miaolingian, Drumian) Marjum Formation of western Utah. Acta Palaeontologica Polonica 66: 663–678. DOI: 10.4202/app.00875.2021

Article featured in press release.


#Yang, Xianfeng, #Julien Kimmig, Dayou Zhai, Yu Liu, Sara R. Kimmig & Shanchi Peng (2021) A juvenile-rich palaeocommunity of the lower Cambrian Chengjiang biota: evidence for palaeo-boom or palaeo-bust? Nature Ecology & Evolution 5: 1082–1088. DOI: 10.1038/s41559-021-01490-4

Cover article.

Article featured in press release.


#Farrell, Úna C., and 103 authors including Julien Kimmig (2021) The Sedimentary Geochemistry and Paleoenvironments Project. Geobiology 19: 545–556. DOI: 10.1111/gbi.12462


#Kimmig, Julien & Paul A. Selden (2021) A new shell-bearing organism from the Cambrian Spence Shale of Utah. Palaeoworld 30: 220–228. DOI: 10.1016/j.palwor.2020.05.003

Article featured in press release.


*,#Leibach, Wade. W, *Nick Rose, Kenneth Bader, Laura J. Mohr, *Kristopher Super & Julien Kimmig (2021) Horseshoe crab trace fossils and associated ichnofauna of the Pony Creek Shale Lagerstätte, Upper Pennsylvanian, Kansas, USA. Ichnos 28: 34–45. DOI: 10.1080/10420940.2020.1811268


#Bicknell, Russell D. C., Patrick Smith, Natalie Schroeder & Julien Kimmig (2020) Reconsidering the ‘phyllocarid’ from the Wade Creek Formation. Alcheringa 44: 481–483. DOI: 10.1080/03115518.2020.1820575


**,#Whitaker, Anna F. & Julien Kimmig (2020) Anthropologically introduced biases in natural history collections, with a case study on the Spence Shale Lagerstätte of Utah. Palaeontologia Electronica 23: a58. DOI: 10.26879/1106


#Lerosey-Aubril, Rudy, Julien Kimmig, Stephen Pates, Jacob Skabelund, Andries Weug & Javier Ortega-Hernández (2020) New exceptionally-preserved panarthropods from the Drumian Wheeler Konservat-Lagerstätte of the House Range of Utah. Papers in Palaeontology 6: 501–531. DOI: 10.1002/spp2.1307


#Yang, Xianfeng, #Julien Kimmig, Bruce S. Lieberman & Shanchi Peng (2020) A new species of the deuterostome Herpetogaster from the early Cambrian Chengjiang biota of South China. The Science of Nature 107: 37. DOI: 10.1007/s00114-020-01695-w


#Kimmig, Julien, *Wade W. Leibach & Bruce S. Lieberman (2020) First occurrence of the problematic vetulicolian Skeemella clavula in the Cambrian Marjum Formation of Utah, USA. Carnets de Géologie 20: 215–221. DOI: 10.4267/2042/70836


**,#Whitaker, Anna, Paul Jamison, James D. Schiffbauer & Julien Kimmig (2020) Re-description of the Spence Shale palaeoscolecids in the light of new morphological features with comments on palaeoscolecid taxonomy and taphonomy. PalZ 94: 661–674­. DOI: 10.1007/s12542-020-00516-9

Article featured in press release.


#Kimmig, Julien, Helena Couto, *Wade W. Leibach & Bruce S. Lieberman (2019) Soft-bodied fossils from the upper Valongo Formation (Middle Ordovician: Dapingian-Darriwilian) of northern Portugal. The Science of Nature 106: 27. DOI: 10.1007/s00114-019-1623-z

Article featured in press release.


#Kimmig, Julien, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff & Bruce S. Lieberman (2019) The Spence Shale Lagerstätte: an Important Window into Cambrian Biodiversity. Journal of the Geological Society of London 176: 609–619. DOI: 10.1144/jgs2018-195

Invited submission.


#Pratt, Brian R. & Julien Kimmig (2019) Extensive burrowing in a middle Cambrian Burgess Shale-type fossil Lagerstätte in northwestern Canada. Geology 47: 231–234. DOI: 10.1130/G45551.1

Article featured in press release.


#Kimmig, Julien, Ronald C. Meyer & Bruce S. Lieberman (2019) Herpetogaster from the early Cambrian of Nevada (Series 2, Stage 4) and its implications for the evolution of deuterostomes. Geological Magazine 156: 172–178. DOI: 10.1017/S0016756818000389


#Seltmann, Katja, and 23 authors including Julien Kimmig (2018) Georeferencing for Research Use (GRU): An integrated geospatial training paradigm for biocollections researchers and data providers. Research Ideas and Outcomes 4: e32449. DOI: 10.3897/rio.4.e32449


#Strotz, Luke C., Erin E. Saupe, Julien Kimmig & Bruce S. Lieberman (2018) Metabolic rates, climate and macroevolution: A case study using Neogene molluscs. Proceedings of the Royal Society B: Biological Sciences 285: 20181292. DOI: 10.1098/rspb.2018.1292

Article featured in news.


#Lieberman, Bruce S. & Julien Kimmig (2018) Museums, Paleontology, and a Biodiversity Science Based Approach. In G. D. Rosenberg, & R. M. Clary (Eds.), Museums at the Forefront of the History and Philosophy of Geology: History Made, History in the Making. Geological Society of America Special Paper 535: 335–348. DOI: 10.1130/20182535(22)


#Strotz, Luke C., Marianna Simões, Matthew Girard, Laura Breitkreuz, Julien Kimmig & Bruce S. Lieberman (2018) Getting somewhere with the Red Queen: Chasing a biologically modern definition of the hypothesis. Biology Letters 14: 20170734. DOI: 10.1098/rslb.2017.0734


#Kimmig, Julien & Brian R. Pratt (2018) Coprolites in the Ravens Throat River Lagerstätte of northwestern Canada: implications for the middle Cambrian food web. Palaios 33: 125–140. DOI: 10.2110/palo.2017.038

Cover article.

Article featured in press release.


#Kimmig, Julien, Luke C. Strotz & Bruce S. Lieberman (2017) The stalked filter feeder Siphusauctum lloydguntheri n. sp. from the middle Cambrian (Series 3: Stage 5) Spence Shale of Utah: Its biological affinities and taphonomy. Journal of Paleontology 91: 902–910. DOI: 10.1017/jpa.2017.57

Article featured in press release.


#Kimmig, Julien & Luke C. Strotz (2017) Coprolites in middle Cambrian (Series 2-3) Burgess Shale-type deposits of Nevada and Utah and their ecological implications. Bulletin of Geosciences 92: 297–309. DOI: 10.3140/bull.geosci.1667


#Lieberman, Bruce S., Richard Kurkewicz, Heather Shinogle, Julien Kimmig & Breandán A. MacGabhann (2017) Disc-shaped fossils resembling porpitids or eldonids from the early Cambrian (Series 2: Stage 4) of western U.S.A. PeerJ 5: e3312. DOI: 10.7717/peerj.3312


#Kimmig, Julien & Brian R. Pratt (2016) Taphonomy of the middle Cambrian (Series 3, Drumian) Ravens Throat River Lagerstätte, Mackenzie Mountains, Northwest Territories, Canada. Lethaia 49: 150–169. DOI: 10.1111/let.12135


#Kimmig, Julien & Brian R. Pratt (2015) Soft-bodied biota from the middle Cambrian (Drumian) Rockslide Formation, Mackenzie Mountains, northwestern Canada. Journal of Paleontology 89: 51–71. DOI: 10.1017/jpa.2014.5


#Kimmig, Julien (2013) Possible secondarily terrestrial lifestyle in the European phytosaur Nicrosaurus kapffi (Late Triassic, Norian) – A preliminary study. In: Triassic Geology and Paleontology: Tanner et al. (Eds.). New Mexico Museum of Natural History and Science, Bulletin 61: 306–312. Link


#Kimmig, Julien & Justin A. Spielmann (2011) Biological factors influencing phytosaur (Archosauria: Phytosauridae) taxonomy: A Prospectus. In: Fossil Record 3: Sullivan et al. (Eds.). New Mexico Museum of Natural History and Science, Bulletin 53: 289–294. Link


#Kimmig, Julien & Gernot Arp (2010) Phytosaur remains from the Norian Arnstadt Formation (Leine Valley, Germany), with reference to European phytosaur habitats. Palaeodiversity 3: 215–224. Link





#Kimmig, Julien (2021) Burgess Shale fauna. The Encyclopedia of Geology, 2nd Edition. Scott Elias & David Alderton (eds.). p. 576–582. DOI: 10.1016/B978-0-12-409548-9.12019-6

Invited submission.


Nicht peer reviewed


Plotnick, Roy E., Brendan M. Anderson, Sandra J. Carlson, Advait M. Jukar, Julien Kimmig & Elizabeth Petsios (2023) Paleontology Is Far More Than New Fossil Discoveries. Scientific American. Link


Kimmig, Julien (2022) Book Review: Plotnick, R. 2022. Explorers of Deep Time: Paleontologists and the History of Life. The Quarterly Review of Biology 97 (4): 310.


Gaines, Robert, Rudy Lerosey-Aubril & Julien Kimmig (2019) Middle Cambrian fossil Lagerstätten of Utah. June 28 – July 02, 2019 Fieldtrip Guide. North American Paleontological Convention 2019: 1–65.  


Kimmig, Julien (2017) Book Review: Suttner, T.J. et al. 2016. Planet Earth—In Deep Time. Palaeozoic Series: Devonian and Carboniferous. The Quarterly Review of Biology 92 (3): 306–307.


Kimmig, Julien (2016) Book Review: Gass, K. 2015. Solving the Mystery of the First Animals on Land: The Fossils of Blackberry Hill. Fossil News: Fall 2016: 51.


Kimmig, Julien (2016) Book Review: Penney, D. and J.E. Jepson. 2014. Fossil Insects: An introduction to palaeoentomology. Priscum: Newsletter of the Paleontological Society: Winter 2016: 42–43.


Eingeladene Vorträge


Kimmig, Julien (2023) Das Kambrium – Ursprung des modernen Lebens. Colloquium, Naturwissenschaftlicher Verein Karlsruhe (Germany), April 2023. November 11th, 2023. Seminar talk.


Kimmig, Julien (2023) Exploring deep time in collections and the field. AGW - Allgemeine Geologie, Karlsruhe Institut für Technologie, Julyth, 2023. Seminar talk.


Kimmig, Julien (2022) The middle Cambrian Spence Shale (Miaolingian: Wuliuan) Lagerstätte: Improving our understanding of a key Cambrian ecosystem. The Pennsylvania State University, March 21st, 2022. Seminar talk.


Kimmig, Julien (2021) Understanding Cambrian diversity through time and space. Friedrich-Alexander-Universität Erlangen-Nürnberg, November 22nd, 2021. Seminar talk.


Kimmig, Julien (2019) Cambrian soft-bodied biotas: Understanding Cambrian ecosystems. Gonzaga University, October 18th, 2019. Seminar talk.


Kimmig, Julien, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff & Bruce S. Lieberman (2019) The middle Cambrian Spence Shale (Miaolingian: Wuliuan) Lagerstätte: Improving our understanding of a key Cambrian ecosystem. North American Paleontological Convention, Riverside, June 23rd, 2019. (KEYNOTE)


Kimmig, Julien (2018) The Cambrian: More than just trilobites. The 57th Annual Kansas City Gem and Mineral Show, March 11th, 2018.


Kimmig, Julien (2018) The Cambrian: More than just trilobites. Science on Tap, January 24th, 2018.


Kimmig, Julien (2017) Cambrian Soft-bodied biotas. Kansas and Missouri Paleontological Society, May 28th, 2017.


Kimmig, Julien (2016) Burgess Shale-type deposits: A window into Cambrian paleoecology. Arkansas Tech University, February 9th, 2016. Seminar talk.

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